Helistra, daughter of Speedwell who is one of four Forestalls who tend the wellbeing of the forests, marries Grimstone, a fellow Forestall. She becomes a mother when their son is born, Tomin.

But when Tomin is being taught the ways of the forest, a knowledge she has been denied, a jealousy arises within Helistra, feeding an evil that has always lurked deep inside her that turns into a hatred that sees her kill Grimstone with his own Rune Staff!

As she strikes, she breaks the rune staff leaving a piece lodged inside of Grimstone. An explosion of white light turns him into stone and makes Helistra disappear. 

Years pass by as Helistra grooms the broken rune staff in the dark evil that has consumed her. Her poisoned mind will not rest until the forests and its keepers are totally destroyed! 

So the paths of conflict and destiny entwine as Speedwell pursues his daughter to make Grimstones broken rune staff whole again to stop her evil, but how and at what cost?